NAGAI Keisuke 永井啓祐

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Nagai Keisuke 永井啓祐

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Researcher Number	30648473
Other IDs
Affiliation (Current)	2025: 名古屋大学, 生物機能開発利用研究センター, 助教
Affiliation (based on the past Project Information) *help	2019 – 2024: 名古屋大学, 生物機能開発利用研究センター, 助教 2016 – 2018: 名古屋大学, 生物機能開発利用研究センター, 特任助教
Review Section/Research Field	Principal Investigator Basic Section 39010:Science in plant genetics and breeding-related / Plant molecular biology/Plant physiology Except Principal Investigator Basic Section 39010:Science in plant genetics and breeding-related
Keywords	Principal Investigator イネ / 洪水耐性 / 浮イネ / 介在分裂組織 / ジベレリン / 節間伸長 / 茎構造 / 低酸素応答 / イネ科作物 / 環境適応 … More / 髄腔 / 節 / 通気組織 / 茎 / 耐水性 / ジベレリン応答性 / 植物生理 / WOX遺伝子 / ストレス / 生理学 / 遺伝学 / 植物 / 撥水性 / 酸素動態 / 植物ホルモン / エチレン / 低酸素 … More Except Principal Investigator イネ / オミクス / 量的形質遺伝子座 / 有用遺伝子同定 Less

有用遺伝子探索を革新するオミクスベースクローニング法の実証
- Principal Investigator
  
  古田智敬
- Project Period (FY)
  2023 – 2026
- Research Category
  
  Grant-in-Aid for Scientific Research (B)
- Review Section
  
  Basic Section 39010:Science in plant genetics and breeding-related
- Research Institution
  Okayama University
Elucidation of a novel water-resistance mechanism in rice stems and construction of a molecular basis for breeding applications.Principal Investigator
- Principal Investigator
  
  永井啓祐
- Project Period (FY)
  2022 – 2025
- Research Category
  
  Grant-in-Aid for Scientific Research (B)
- Review Section
  
  Basic Section 39010:Science in plant genetics and breeding-related
- Research Institution
  Nagoya University
Elucidation of molecular mechanism of water stress tolerance in rice internode.Principal Investigator
- Principal Investigator
  
  Keisuke Nagai
- Project Period (FY)
  2019 – 2021
- Research Category
  
  Grant-in-Aid for Early-Career Scientists
- Review Section
  
  Basic Section 39010:Science in plant genetics and breeding-related
- Research Institution
  Nagoya University
Elucidation of molecular mechanism of flood tolerance by internode elongation in deepwater ricePrincipal Investigator
- Principal Investigator
  
  Nagai Keisuke
- Project Period (FY)
  2016 – 2018
- Research Category
  
  Grant-in-Aid for Young Scientists (B)
- Research Field
  
  Plant molecular biology/Plant physiology
- Research Institution
  Nagoya University

All 2023 2022 2021 2020 2019 2018 2017 2016

All Journal Article Presentation

[Journal Article] Regulator of Awn Elongation 3, an E3 ubiquitin ligase, is responsible for loss of awns during African rice domestication2023
- Author(s)
  Bessho-Uehara Kanako、Masuda Kengo、Wang Diane R.、Angeles-Shim Rosalyn B.、Obara Keisuke、Nagai Keisuke、Murase Riri、Aoki Shin-ichiro、Furuta Tomoyuki、Miura Kotaro、Wu Jianzhong、Yamagata Yoshiyuki、Yasui Hideshi、Kantar Michael B.、Yoshimura Atsushi、Kamura Takumi、McCouch Susan R.、Ashikari Motoyuki
- Journal Title
  
  Proceedings of the National Academy of Sciences
  
  Volume: 120 Issue: 4
- DOI
  10.1073/pnas.2207105120
- Peer Reviewed / Open Access / Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-20H03208, KAKENHI-PROJECT-20K22644, KAKENHI-PROJECT-23K23575, KAKENHI-PROJECT-21K15115
[Journal Article] Can the Wild Perennial, Rhizomatous Rice Species Oryza longistaminata be a Candidate for De Novo Domestication?2023
- Author(s)
  Tong Shuai、Ashikari Motoyuki、Nagai Keisuke、Pedersen Ole
- Journal Title
  
  Rice
  
  Volume: 16 Issue: 1
- DOI
  10.1186/s12284-023-00630-7
- Peer Reviewed / Open Access / Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-23K23575
[Journal Article] Molecular mechanism of internode elongation in rice2023
- Author(s)
  Nagai Keisuke、Ashikari Motoyuki
- Journal Title
  
  Breeding Science
  
  Volume: 73 Issue: 2 Pages: 108-116
- DOI
  10.1270/jsbbs.22086
- ISSN
  1344-7610, 1347-3735
- Language
  English
- Peer Reviewed / Open Access
- Data Source
  KAKENHI-PROJECT-23K23575
[Journal Article] SNORKEL Genes Relating to Flood Tolerance Were Pseudogenized in Normal Cultivated Rice2022
- Author(s)
  Nagai Keisuke、Kurokawa Yusuke、Mori Yoshinao、Minami Anzu、Reuscher Stefan、Wu Jianzhong、Matsumoto Takashi、Ashikari Motoyuki
- Journal Title
  
  Plants
  
  Volume: 11 Issue: 3 Pages: 376-376
- DOI
  10.3390/plants11030376
- Peer Reviewed / Open Access / Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-19K15815, KAKENHI-PROJECT-22K14880, KAKENHI-PLANNED-20H05912, KAKENHI-PROJECT-22H04978
[Journal Article] Deep Fluorescence Observation in Rice Shoots via Clearing Technology2022
- Author(s)
  Niimi Yoko、Nagai Keisuke、Ashikari Motoyuki、Mizuta Yoko
- Journal Title
  
  Journal of Visualized Experiments
  
  Volume: 184 Issue: 184 Pages: 1-13
- DOI
  10.3791/64116
- Peer Reviewed / Open Access
- Data Source
  KAKENHI-ORGANIZER-20H05778, KAKENHI-PLANNED-20H05779, KAKENHI-PROJECT-23K23575, KAKENHI-PLANNED-20H05912, KAKENHI-PROJECT-22H04978
[Journal Article] Elucidation of molecular mechanism of internode elongation in rice2022
- Author(s)
  Nagai Keisuke
- Journal Title
  
  Breeding Research
  
  Volume: 24 Issue: 2 Pages: 195-201
- DOI
  10.1270/jsbbr.22J09
- ISSN
  1344-7629, 1348-1290
- Year and Date
  2022-12-01
- Language
  Japanese
- Peer Reviewed / Open Access
- Data Source
  KAKENHI-PROJECT-23K23575
[Journal Article] Noninvasive imaging of hollow structures and gas movement revealed the gas partial‐pressure‐gradient‐driven long‐distance gas movement in the aerenchyma along the leaf blade to submerged organs in rice2021
- Author(s)
  Yin Yong‐Gen、Mori Yoshinao、Suzui Nobuo、Kurita Keisuke、Yamaguchi Mitsutaka、Miyoshi Yuta、Nagao Yuto、Ashikari Motoyuki、Nagai Keisuke、Kawachi Naoki
- Journal Title
  
  New Phytologist
  
  Volume: 232 Issue: 5 Pages: 1974-1984
- DOI
  10.1111/nph.17726
- Peer Reviewed / Open Access
- Data Source
  KAKENHI-PROJECT-19K15815, KAKENHI-PLANNED-20H05912, KAKENHI-PROJECT-20H04465
[Journal Article] Antagonistic regulation of the gibberellic acid response during stem growth in rice2020
- Author(s)
  Nagai Keisuke et al.
- Journal Title
  
  Nature
  
  Volume: 584 Issue: 7819 Pages: 109-114
- DOI
  10.1038/s41586-020-2501-8
- Peer Reviewed / Open Access / Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-20K06043, KAKENHI-PROJECT-19K15815, KAKENHI-PROJECT-17H03751, KAKENHI-PLANNED-16H06466, KAKENHI-PLANNED-17H06471, KAKENHI-PROJECT-20H00424, KAKENHI-PLANNED-17H06473, KAKENHI-PROJECT-19H02930
[Journal Article] Metabolite and Phytohormone Profiling Illustrates Metabolic Reprogramming as an Escape Strategy of Deepwater Rice during Partially Submerged Stress2020
- Author(s)
  Fukushima Atsushi、Kuroha Takeshi、Nagai Keisuke、Hattori Yoko、Kobayashi Makoto、Nishizawa Tomoko、Kojima Mikiko、Utsumi Yoshinori、Oikawa Akira、Seki Motoaki、Sakakibara Hitoshi、Saito Kazuki、Ashikari Motoyuki、Kusano Miyako
- Journal Title
  
  Metabolites
  
  Volume: 10 Issue: 2 Pages: 68-68
- DOI
  10.3390/metabo10020068
- NAID
  120007132876
- Peer Reviewed / Open Access
- Data Source
  KAKENHI-PROJECT-18K06274, KAKENHI-PROJECT-19K15815, KAKENHI-PROJECT-17K07663
[Journal Article] Evaluation of backcrossed pyramiding lines of the yield-related gene and the bacterial leaf blight resistant genes2020
- Author(s)
  Shuto Yamada, Yusuke Kurokawa, Keisuke Nagai, Rosalyn B Angeles-Shim, Hideshi Yasui, Naruto Furuya, Atsushi Yoshimura, Kazuyuki Doi, Motoyuki Ashikari, and Hidehiko Sunohara
- Journal Title
  
  Journal of international cooperation for agricultural development
  
  Volume: 18 Pages: 18-28
- NAID
  120007182024
- Peer Reviewed / Open Access / Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-19K15815
[Journal Article] Diel O2 dynamics in partially and completely submerged deepwater rice: Leaf gas films enhance internodal O2 status, influence gene expression and accelerate stem elongation for ‘Snorkelling’ during submergence.2019
- Author(s)
  Y. Mori, Y. Kurokawa, M. Koike. A. I. Malik, T. D. Colmer, M. Ashikari, O. Pedersen and K. Nagai
- Journal Title
  
  Plant and Cell Physiology
  
  Volume: 60 Issue: 5 Pages: 973-985
- DOI
  10.1093/pcp/pcz009
- Peer Reviewed / Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-19K15815, KAKENHI-PLANNED-17H06473
[Journal Article] Breeding Applications and Molecular Basis of Semi-dwarfism in Rice2018
- Author(s)
  Nagai Keisuke、Hirano Ko、Angeles-Shim Rosalyn B.、Ashikari Motoyuki
- Journal Title
  
  Rice Genomics, Genetics and Breeding
  
  Volume: - Pages: 155-176
- DOI
  10.1007/978-981-10-7461-5_9
- ISBN
  9789811074608, 9789811074615
- Peer Reviewed
- Data Source
  KAKENHI-PROJECT-16K18565
[Journal Article] Time-Course Transcriptomics Analysis Reveals Key Responses of Submerged Deepwater Rice to Flooding2018
- Author(s)
  Minami Anzu、Yano Kenji、Gamuyao Rico、Nagai Keisuke et. al.
- Journal Title
  
  Plant Physiology
  
  Volume: 176 Issue: 4 Pages: 3081-3102
- DOI
  10.1104/pp.17.00858
- Peer Reviewed / Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-16K18565, KAKENHI-PROJECT-17K15209, KAKENHI-PLANNED-17H06473, KAKENHI-PROJECT-16J08722, KAKENHI-PROJECT-18K06274, KAKENHI-PROJECT-17K15136
[Journal Article] Rice leaf hydrophobicity and gas films are conferred by a wax synthesis gene (LGF1) and contribute to flood tolerance.2018
- Author(s)
  Kurokawa, Y., Nagai, K., Huan, P. D., Shimazaki, K., Qu, H., Mori, Y., Toda, Y., Kuroha, T., Hayashi, N., Aiga, S., Itoh, J.-i., Yoshimura, A., Sasaki-Sekimoto, Y., Ohta, H., Shimojima, M., Malik, A. I., Pedersen, O., Colmer, T. D. and Ashikari, M.
- Journal Title
  
  New Phytologist
  
  Volume: － Issue: 4 Pages: 1558-1569
- DOI
  10.1111/nph.15070
- Peer Reviewed / Open Access / Int'l Joint Research
- Data Source
  KAKENHI-PUBLICLY-16H01230, KAKENHI-PLANNED-17H06473, KAKENHI-PROJECT-16H04857, KAKENHI-PROJECT-18K06274, KAKENHI-PROJECT-16K18565
[Journal Article] Ethylene-gibberellin signaling underlies adaptation of rice to periodic flooding2018
- Author(s)
  T Kuroha, K Nagai, R Gamuyao, D R Wang, T Furuta, M Nakamori, T Kitaoka, K Adachi, A Minami, Y Mori, K Mashiguchi, Y Seto, S Yamaguchi, M Kojima, H Sakakibara, J Wu, K Ebana, N Mitsuda, M Ohme-Takagi, S Yanagisawa, M Yamasaki, R Yokoyama, K Nishitani, T Mochizuki, G Tamiya, S R McCouch, and M Ashikari
- Journal Title
  
  Science
  
  Volume: 361 Issue: 6398 Pages: 181-186
- DOI
  10.1126/science.aat1577
- NAID
  120006529624
- Peer Reviewed / Open Access / Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-18K06274, KAKENHI-PROJECT-17K19374, KAKENHI-PLANNED-17H06473, KAKENHI-PLANNED-17H06474, KAKENHI-PLANNED-18H05489, KAKENHI-PROJECT-17H03691, KAKENHI-PROJECT-16K07389, KAKENHI-PROJECT-16K08638, KAKENHI-PROJECT-16K18565, KAKENHI-PROJECT-17K15136
[Journal Article] Hormone Distribution And Transcriptome Profiles In Bamboo Shoots Provide Insights On Bamboo Stem Emergence And Growth2017
- Author(s)
  Rico Gamuyao, Keisuke Nagai, Madoka Ayano, Yoshinao Mori, Anzu Minami, Mikiko Kojima, Takamasa Suzuki, Hitoshi Sakakibara, Tetsuya Higashiyama, Motoyuki Ashikari, Stefan Reuscher
- Journal Title
  
  PLANT & CELL PHYSIOLOGY
  
  Volume: 58 Issue: 4 Pages: 702-716
- DOI
  10.1093/pcp/pcx023
- NAID
  120006343684
- Peer Reviewed / Open Access / Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-16K18565
[Journal Article] eQTLs Regulating Transcript Variations Associated with Rapid Internode Elongation in Deepwater Rice2017
- Author(s)
  1.Takeshi Kuroha, Keisuke Nagai, Yusuke Kurokawa, Yoshiaki Nagamura, Miyako Kusano, Hideshi Yasui, Motoyuki Ashikari, Atsushi Fukushima
- Journal Title
  
  Frontiers in Plant Science
  
  Volume: 8 Pages: 1753-1753
- DOI
  10.3389/fpls.2017.01753
- NAID
  120007134740
- Peer Reviewed / Open Access
- Data Source
  KAKENHI-PROJECT-17K07663, KAKENHI-PROJECT-16K18565, KAKENHI-PLANNED-17H06473
[Presentation] イネ節間伸長の分子機構の解明2022
- Author(s)
  永井啓祐
- Organizer
  日本育種学会第141回講演会
- Invited
- Data Source
  KAKENHI-PROJECT-19K15815
[Presentation] Elucidation of water-tolerant mechanism by gas permeability in rice node2022
- Author(s)
  Keisuke Nagai
- Organizer
  International Society of Plant Anaerobiosis Conference
- Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-23K23575
[Presentation] 節間伸長促進因子SNORKEL-LIKEsの偽遺伝子化による水田イネ短稈化機構の解明2022
- Author(s)
  永井啓祐、芦苅基行
- Organizer
  日本育種学会第141回講演会
- Data Source
  KAKENHI-PROJECT-19K15815
[Presentation] The regulation of phase transition in rice internode2021
- Author(s)
  永井啓祐、森欣順、石川慎、Rico Gamuyao、新美陽子、保浦徳典、福田萌莉、榊原均、古田智敬、久野裕、佐藤和広、赤木剛士、小嶋美紀子、竹林　裕美子、福島敦史、氷室泰代、小林正智、呉健忠、アキリ亘、吉田綾、辻寛之、佐藤豊、芦苅基行
- Organizer
  第62回日本植物生理学会年会
- Data Source
  KAKENHI-PROJECT-19K15815
[Presentation] イネ節間におけるジベレリンに応答した拮抗的伸長制御機構2021
- Author(s)
  永井啓祐、森欣順、石川慎、Rico Gamuyao、新美陽子、保浦徳典、福田萌莉、榊原均、古田智敬、久野裕、佐藤和広、赤木剛士、小嶋美紀子、竹林　裕美子、福島敦史、氷室泰代、小林正智、呉健忠、アキリ亘、吉田綾、辻寛之、佐藤豊、芦苅基行
- Organizer
  日本育種学会第139回講演会
- Data Source
  KAKENHI-PROJECT-19K15815
[Presentation] ACE1とDEC１によるイネ節間伸長のantagonistic制御22020
- Author(s)
  永井啓祐、森欣順、石川慎、Rico Gamuyao、新美陽子、保浦徳典、福田萌莉、榊原均、古田智敬、久野裕、佐藤和広、赤木剛士、小嶋美紀子、竹林　裕美子、福島敦史、氷室泰代、小林正智、呉健忠、アキリ亘、吉田綾、辻寛之、佐藤豊、芦苅基行
- Organizer
  日本植物学会第84回大会
- Data Source
  KAKENHI-PROJECT-19K15815
[Presentation] イネ節間伸長の開始制御機構の解明2020
- Author(s)
  永井啓祐、森欣順、石川慎、Rico Gamuyao、新美陽子、保浦徳典、福田萌莉、榊原均、古田智敬、久野裕、佐藤和広、赤木剛士、小嶋美紀子、竹林　裕美子、福島敦史、氷室泰代、小林正智、呉健忠、アキリ亘、吉田綾、辻寛之、佐藤豊、芦苅基行
- Organizer
  日本育種学会第138回講演会
- Data Source
  KAKENHI-PROJECT-19K15815
[Presentation] イネ節間伸長における拮抗的制御機構2020
- Author(s)
  永井啓祐，芦苅基行
- Organizer
  第61回日本植物生理学会年会
- Data Source
  KAKENHI-PROJECT-19K15815
[Presentation] Antagonistic regulatory mechanism by accelerating and decelerating factors in internode elongation of rice for flooding adaptation.2019
- Author(s)
  Keisuke Nagai, Shin Ishikawa, Mori Yoshinao, Yoko Niimi, Tokunori Hobo, Motoyuki Ashikari
- Organizer
  The 13th International Society of Plant Anaerobiosis Conference
- Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-19K15815
[Presentation] イネ節の効率的な気体透過性による耐水性機構の解明2018
- Author(s)
  永井啓祐
- Organizer
  第34回資源植物科学シンポジウム、第10回植物ストレス科学研究シンポジウム
- Invited
- Data Source
  KAKENHI-PROJECT-16K18565
[Presentation] イネが水田で生きるためには～コンペイ糖状細胞の発見とガス交換の仕組み～2017
- Author(s)
  永井啓祐
- Organizer
  2017年遺伝学研究所研究会「イネ分子遺伝学の方向性」
- Data Source
  KAKENHI-PROJECT-16K18565
[Presentation] 植物が水辺で生きる2017
- Author(s)
  永井啓祐
- Organizer
  日本育種学会第132回講演会
- Invited
- Data Source
  KAKENHI-PROJECT-16K18565
[Presentation] イネがイネであるために～茎の形態から～2017
- Author(s)
  永井啓祐
- Organizer
  第3回農学中手の会
- Data Source
  KAKENHI-PROJECT-16K18565
[Presentation] Oxygen transport through rice node tissue2017
- Author(s)
  Keisuke Nagai
- Organizer
  IGER symposium on Long0distance signaling in plant
- Data Source
  KAKENHI-PROJECT-16K18565
[Presentation] Anatomic analysis of aerenchyma in grasses2016
- Author(s)
  Keisuke Nagai, Yoshinao Mori, Tim Colmer, Ole Pedersen and Motoyuki Ashikari
- Organizer
  International society for plant anaerobiosis
- Place of Presentation
  Copenhagen, Denmark
- Year and Date
  2016-09-05
- Int'l Joint Research
- Data Source
  KAKENHI-PROJECT-16K18565
[Presentation] イネ節間伸長制御機構の組織化学的解析2016
- Author(s)
  大里美咲、永井啓祐、芦苅基行
- Organizer
  日本育種学会
- Place of Presentation
  鳥取大学
- Year and Date
  2016-09-25
- Data Source
  KAKENHI-PROJECT-16K18565
[Presentation] イネ科植物における節間通気組織の形態学的解析2016
- Author(s)
  永井啓祐、大里美咲、芦苅基行
- Organizer
  イネ遺伝学・分子生物学ワークショップ
- Place of Presentation
  名古屋大学
- Year and Date
  2016-07-04
- Data Source
  KAKENHI-PROJECT-16K18565

1. 古田智敬 (70774008)

# of Collaborated Projects: 1 results

# of Collaborated Products: 0 results
2. 鳥羽大陽 (10585160)

# of Collaborated Projects: 1 results

# of Collaborated Products: 0 results
3. 黒羽剛 (50415155)

# of Collaborated Projects: 1 results

# of Collaborated Products: 2 results
4. FUKUSHIMA Atsushi

# of Collaborated Projects: 0 results

# of Collaborated Products: 1 results
5. 嘉村巧

# of Collaborated Projects: 0 results

# of Collaborated Products: 1 results
6. 伊藤純一

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# of Collaborated Products: 1 results
7. 水多陽子

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# of Collaborated Products: 1 results

NAGAI Keisuke 永井 啓祐

Nagai Keisuke 永井 啓祐

Research Projects

Research Products

Co-Researchers

有用遺伝子探索を革新するオミクスベースクローニング法の実証

Principal Investigator

Project Period (FY)

Research Category

Review Section

Research Institution

Elucidation of a novel water-resistance mechanism in rice stems and construction of a molecular basis for breeding applications.Principal Investigator

Principal Investigator

Project Period (FY)

Research Category

Review Section

Research Institution

Elucidation of molecular mechanism of water stress tolerance in rice internode.Principal Investigator

Principal Investigator

Project Period (FY)

Research Category

Review Section

Research Institution

Elucidation of molecular mechanism of flood tolerance by internode elongation in deepwater ricePrincipal Investigator

Principal Investigator

Project Period (FY)

Research Category

Research Field

Research Institution

[Journal Article] Regulator of Awn Elongation 3, an E3 ubiquitin ligase, is responsible for loss of awns during African rice domestication2023

Author(s)

Journal Title

DOI

Data Source

[Journal Article] Can the Wild Perennial, Rhizomatous Rice Species Oryza longistaminata be a Candidate for De Novo Domestication?2023

Author(s)

Journal Title

DOI

Data Source

[Journal Article] Molecular mechanism of internode elongation in rice2023

Author(s)

Journal Title

DOI

ISSN

Language

Data Source

[Journal Article] SNORKEL Genes Relating to Flood Tolerance Were Pseudogenized in Normal Cultivated Rice2022

Author(s)

Journal Title

DOI

Data Source

[Journal Article] Deep Fluorescence Observation in Rice Shoots via Clearing Technology2022

Author(s)

Journal Title

DOI

Data Source

[Journal Article] Elucidation of molecular mechanism of internode elongation in rice2022

Author(s)

Journal Title

DOI

ISSN

Year and Date

Language

Data Source

[Journal Article] Noninvasive imaging of hollow structures and gas movement revealed the gas partial‐pressure‐gradient‐driven long‐distance gas movement in the aerenchyma along the leaf blade to submerged organs in rice2021

Author(s)

Journal Title

DOI

Data Source

[Journal Article] Antagonistic regulation of the gibberellic acid response during stem growth in rice2020

Author(s)

Journal Title

DOI

Data Source

[Journal Article] Metabolite and Phytohormone Profiling Illustrates Metabolic Reprogramming as an Escape Strategy of Deepwater Rice during Partially Submerged Stress2020

Author(s)

Journal Title

DOI

NAID

Data Source

NAGAI Keisuke 永井啓祐

Nagai Keisuke 永井啓祐